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This American geneticist discovered that a chromosome pair determines an organism's sex.
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This American psychologist pioneered the study of operant conditioning using pigeons and rats.
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This German-American biophysicist showed that bacteria evolve resistance through random mutation, not adaptation.
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This Japanese immunologist explained how the body generates a vast diversity of antibodies.
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This statistics-minded nursing pioneer, first name _ _ _ _ _ _ _ _, transformed hospital sanitation practices.
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A biologist named Dobzhansky famously wrote that nothing in biology makes sense except in light of _ _ _ _ _ _ _ _ _.
EVOLUTIONThis U.S. Army physician proved that mosquitoes, not direct contact, spread yellow fever.
REEDThis American botanist and horticulturist was among the first in the U.S. to recognize the importance of Mendel's work.
BAILEYThis FDA reviewer blocked U.S. approval of thalidomide, preventing a wave of birth defects.
KELSEYThis French physician is credited with founding humane, scientific treatment of the mentally ill.
PINELThis British developmental biologist showed that mature cell nuclei can be reprogrammed to grow a new organism.
GURDONThis Nobel-winning geneticist, first name _ _ _ _ _ _ _, discovered mobile genetic elements in corn.
BARBARAThis American biologist led the team that first sequenced the complete genome of a free-living organism.
VENTERThis British naturalist proposed evolution by natural selection after his voyage on HMS Beagle.
DARWINThis French chemist and microbiologist disproved spontaneous generation and developed the germ theory of disease.
PASTEURThis Australian pathologist co-discovered the ulcer-causing bacterium Helicobacter pylori.
WARRENThis American paleontologist's fossil finds revealed that some dinosaurs cared for their young in nests.
HORNERThis American chemical engineer pioneered directed evolution to design new enzymes in the lab.
ARNOLDThis American primatologist devoted decades to studying mountain gorillas in Rwanda before her murder in 1985.
FOSSEYThis Austrian zoologist decoded the 'waggle dance' honeybees use to tell hivemates where to find food.
FRISCHThis Japanese scientist discovered how to reprogram adult cells into induced pluripotent stem cells.
YAMANAKAThis American microbiologist discovered Archaea, redrawing the tree of life into three domains.
WOESEThis American biologist explained the chemistry behind how the eye detects light and color.
WALDThis British naturalist collected roughly 30,000 plant specimens while sailing with Captain Cook.
BANKSThis Swedish botanist created the two-part naming system still used to classify every living species.
LINNAEUSThis microbiologist discovered streptomycin, the first effective antibiotic treatment for tuberculosis.
WAKSMANThis British primatologist redefined the human-animal boundary by documenting tool use among chimpanzees.
GOODALLThis American biochemist pioneered recombinant DNA technology by splicing genes from different organisms.
BERGThis English scientist coined the term 'cell' after examining cork under a microscope in 1665.
HOOKEThis South African-born biologist established a tiny roundworm as a model organism for genetics research.
BRENNERThis French naval officer co-invented the Aqua-Lung and became a pioneering ocean documentarian.
COUSTEAUThis American paleontologist proposed 'punctuated equilibrium' as a pattern for how evolution proceeds.
GOULDThis German physician coined the term 'magic bullet' and pioneered chemotherapy for infectious disease.
EHRLICHNew doctors still swear a version of the Hippocratic _ _ _ _, attributed to an ancient Greek physician.
OATHThis American biochemist helped decipher how RNA codons correspond to specific amino acids.
NIRENBERGThis American animal scientist redesigned livestock handling facilities by drawing on her own autism.
GRANDINThis Dutch-British zoologist helped found the modern scientific study of animal behavior.
TINBERGENThis German scientist co-proposed that the cell is the basic structural unit of all living things.
SCHWANNThis British evolutionary biologist popularized gene-centered evolution in his book The Selfish Gene.
DAWKINSThis British chemist's X-ray images were crucial evidence for the structure of DNA.
FRANKLINThis Indian-American biochemist helped crack the genetic code and later synthesized the first artificial gene.
KHORANAThis French bacteriologist proved that body lice transmit the disease typhus.
NICOLLEThis Australian neurophysiologist proved that signals cross the gap between neurons chemically, not electrically.
ECCLESThis English physician pioneered vaccination after showing that cowpox exposure protects against smallpox.
JENNERThis Scottish bacteriologist accidentally discovered penicillin in a contaminated petri dish in 1928.
FLEMINGThis Australian physician proved that a bacterium, not stress, causes most stomach ulcers.
MARSHALLThis American agronomist's high-yield wheat varieties are credited with saving a billion lives from famine.
BORLAUGThis Austrian biochemist's base-pairing rules helped Watson and Crick decode DNA's structure.
CHARGAFFThis British geneticist shared a Nobel Prize for discovering the proteins that control the cell division cycle.
NURSEThis British naturalist independently proposed natural selection at nearly the same time as Darwin.
WALLACEThis Flemish anatomist's detailed 1543 dissections corrected centuries of anatomical errors.
VESALIUSThis American physiologist founded the first pharmacology department at a U.S. medical school.
ABELThis Canadian neurosurgeon mapped brain functions by electrically stimulating patients during surgery.
PENFIELDThis Canadian-American molecular biologist shared a Nobel Prize for discovering catalytic properties of RNA.
ALTMANThis British statistician united Mendelian genetics with Darwinian evolution in the 1930s.
FISHERThis Russian physiologist discovered classical conditioning while studying the salivation of dogs.
PAVLOVThis Australian pharmacologist turned penicillin from a lab curiosity into a mass-produced wartime drug.
FLOREYThis English physician demonstrated that blood circulates continuously around the body, pumped by the heart.
HARVEYThis early American botanist was called by Linnaeus the 'greatest natural botanist in the world.'
BARTRAMThis American molecular biologist shared the Nobel Prize for co-discovering telomerase.
GREIDERThis American biochemist co-developed CRISPR gene editing into a precise, programmable tool.
DOUDNAThis American chemist discovered that RNA molecules can catalyze reactions on their own.
CECHThis American biologist's 1962 book Silent Spring helped launch the modern environmental movement.
CARSONThis Dutch tradesman and lens-grinder, often called the father of microbiology, was named _ _ _ _ _ _ _ van Leeuwenhoek.
ANTONIEThis American virologist developed a rival oral polio vaccine using a weakened live virus.
SABINThis British physicist's X-ray diffraction images of DNA supported the double-helix model.
WILKINSThis American agricultural scientist promoted peanuts and sweet potatoes as soil-restoring alternatives to cotton.
CARVERThis German physician identified the specific bacteria that cause tuberculosis and cholera.
KOCHThis American microbiologist showed in 1944 that DNA, not protein, carries hereditary information between generations.
AVERYThis ornithologist conducted pioneering systematic bird surveys across India for decades.
ALIThis German zoologist coined the word 'ecology' and championed the idea that embryos replay evolutionary history.
HAECKELThis physician became the first woman to earn a medical degree in the United States, in 1849.
BLACKWELLThis British scientist co-discovered the double-helix structure of DNA in 1953 alongside James Watson.
CRICKThis French naturalist proposed an early, ultimately incorrect theory of inherited acquired traits.
LAMARCKThis American virologist developed the first effective injectable polio vaccine in 1955.
SALKThis American scientist co-discovered DNA's double-helix structure alongside Francis Crick.
WATSONThis American biologist founded the field of sociobiology after decades studying ants.
WILSONThis American biochemist studied the protein machinery that copies chromosomes before cell division.
ALBERTSThis American molecular biologist co-discovered telomerase, the enzyme that protects chromosome ends.
BLACKBURNThis American neuroscientist helped explain how olfactory receptors let us distinguish thousands of odors.
BUCKThis Spanish histologist established that the nervous system is made of individual cells, not a continuous web.
CAJALThis scientist, first name _ _ _ _ _ _, founded the field of microbial ecology while working in Russia and France.
SERGEIThis Swiss-American naturalist pioneered the study of fish classification and glacial geology.
AGASSIZThis Italian neurobiologist, first name _ _ _ _, discovered nerve growth factor and won a Nobel Prize.
RITAThis American-French naturalist painted and catalogued hundreds of North American bird species.
AUDUBONThis Austrian zoologist studied imprinting in geese, who would follow him as if he were their mother.
LORENZThis German pathologist declared that all cells arise from pre-existing cells.
VIRCHOWThis British embryologist led the team that cloned Dolly the sheep in 1996.
WILMUTThis Italian scientist's 1668 experiments with sealed jars helped disprove spontaneous generation of maggots.
REDIThis Austrian monk's pea-plant experiments in a monastery garden laid the foundation for modern genetics.
MENDELThis Scottish pharmacologist developed beta-blockers, revolutionizing treatment for heart disease.
BLACKThis German physiologist developed the first antitoxin treatment for diphtheria, winning the first Nobel in Medicine.
BEHRINGNicknamed 'Darwin's Bulldog,' this British biologist fiercely defended evolutionary theory in public debates.
HUXLEYThis American biologist proposed that mitochondria and chloroplasts evolved from engulfed bacteria.
MARGULISThis American microbiologist developed dozens of vaccines, including those for measles and mumps.
HILLEMANThis American geneticist's fruit-fly experiments proved that genes are carried on chromosomes.
MORGANThis scientist's discovery of the ABO _ _ _ _ _ group system made safe transfusions possible for the first time.
BLOODThis British physician proved that mosquitoes transmit the parasite that causes malaria.
ROSSThis German-American biologist defined a species as a group that can interbreed and produce fertile offspring.
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